From aceska@telus.net Fri Nov 18 00:24:53 2005 From: aceska@telus.net (Adolf Ceska) Date: Thu, 17 Nov 2005 16:24:53 -0800 Subject: [BEN-L]BEN # 352 Message-ID: <000001c5ebd6$83ead0a0$f1dcb440@HPLAPTOP001> BBBBB EEEEEE NN N ISSN 1188-603X BB B EE NNN N BBBBB EEEEE NN N N BOTANICAL BB B EE NN NN ELECTRONIC BBBBB EEEEEE NN N NEWS No. 352 November 17, 2005 aceska@telus.net Victoria, B.C. ----------------------------------------------------------- Dr. A. Ceska, P.O.Box 8546, Victoria, B.C. Canada V8W 3S2 ----------------------------------------------------------- BEN issues 352 and 353 are dedicated to the memory of BORIS ALEXANDROVICH YURTSEV who died in St. Petersburg on December 14, 2004 BORIS ALEXANDROVICH YURTSEV (1932-2004) Boris Alexandrovich Yurtsev was born on March 15, 1932 in the very center of Moscow on Arbat Street, now famous for its artists and street vendors. Three institutions that were nearby played essential roles in the life of Boris Yurtsev: his high school, the Geobotanical Department of the Moscow State University, and the Moscow Conservatory of Music. At the Conservatory, young Boris could be found "eight times a week". >From that point on, music and poetry were important parts of his life. During WWII, Boris Alexandrovich was evacuated from Moscow to Kazan, and his first field trips to the outdoors awakened his botanical interests. After his return to Moscow in 1944, Boris Alexandrovich joined the circle of young naturalists who regularly visited the Geobotanical Department of Moscow State University. He helped to set up the exposition that showcased the diversity of the flora of the Soviet Union. Under the direction of Ivan Serebryakov, Boris Alexandrovich investigated the periodicity of the underground biomass in grassy oak-forests, and this study resulted in his first scientific publication. This was not unusual in the circle of talented students of which he was a part. At that time he also found a tutor in amateur botanist B.M. Kul'kov, with whom he made numerous field trips in the Moscow area. In 1950 Boris Alexandrovich entered the Faculty of Biology and Soil Science of Moscow State University. There were several other bright students in his class, and Boris Alexandrovich studied with them the ecology of the southern boundary of fir (_Abies alba_) in the Moscow area. Field trips he made in the early 1950s triggered his interest in phytogeography and florogenesis of larger floristic areas: in 1952 he was in the Crimea, in 1953 in Tjan-Shan, in 1954 in Tadjikistan and Azerbaijan. His degree thesis (a rough equivalent of M.Sc.) dealt with the evolution of certain forms of one section of maples (genus _Acer_). In 1955, Prof. T.A. Rabotnov recommended his favourite student to Prof. B.A. Tikhomirov, who at that time was setting up a group to study the Russian Arctic. Yurtsev joined the group that was based in the main botanical institute of the Soviet Union, BIN, in Leningrad, and participated in many field trips to the Russian Arctic. One year later, Prof. A.I. Tolmachev was starting his monumental 10-volume treatise _Flora of the Russian Arctic_, now partly available in English translation: B.A. Yurtzev was initially assigned to work on the massive family Leguminosae; however, his role in the project grew, and after the death of A.I. Tolmachev, Yurtzev completed that treatise in 1987. With Prof. Tolmachev, Boris Alexandrovich took part in the botanical and pedological expedition to the lower Lena basin. This expedition greatly contributed to the knowledge of northern flora and the cytotaxonomy of northern plants. In 1958, Boris Alexandrovich spent the summer in the northern Yakutsk region. This study resulted in his doctoral dissertation _Botanical and geographical analysis of the flora and vegetation of Suntar-Khayat Range_. This was the first attempt of multidisciplinary analysis of the montane floras of North- Eastern Asia. The principal chapter of this dissertation was later published in 1968, in the classical monograph _Flora of Suntar-Khayat: Aspects of the history of high mountain landscapes of North-Eastern Asia_. With B.A. Tolmachev, Boris Alexandrovich wrote an important essay entitled _The Evolution of the Arctic flora and its ties to the history of the North Sea_ (1970). In it, a concept of the early (Pliocene) origin of the Asian taiga flora was further advanced. Later, Yurtzev developed a hypothesis of the existence of dry and cold "tundra-steppe" in the Pleistocene of North-East Asia, backed by the discovery of what he believed were the extant pockets of relic steppe-like grass communities in dry continental portions of Yakutia. Therefore, Yurtzev was among the first biologists who speculated that in the Pleistocene and early Holocene, ecosystems existed that do not have analogs in the modern world. This notion has since been widely supported by paleobotanical data. Boris Alexandrovich Yurtsev had vast experience with the flora of the Chukotka Peninsula, and he also made many field trips to arctic and subarctic regions of Yakutsia (1974- 1976), northernmost regions of the Ural Mountains (1996-1998) and to Mongolia (1988). Thanks to his close friend Dave Murray (University of Alaska in Fairbanks) he visited Alaska several times (first in 1980). Later he also botanized in the Canadian Arctic regions (1990, 1999), Svalbard (Spitsbergen) and northern Norway (1995), Central Alps (1994), Greenland (1997), and Lapland (1996). Yurtsev's interest in the northern flora and the flora of the Russian Far East led him to the study of the issue of the Bering Bridge and Beringia and to the idea of the Panarctic Flora, an attempt to deal with the Holarctic region as a whole. This led to the initiation of the Panarctic Flora and his cooperation with David Murray (USA), George Packer (Canada), and Olaf Gjaerevoll (Norway). Boris Alexandrovich Yurtsev was an excellent field botanist. He described about 170 new species and subspecies of flowering plants, and one new genus _Claytoniella_ (Portulacaceae): http://herba.msu.ru/russian/journals/herba/yurtsev/yurtsev_taxa.html Several taxa were named in his honour. A famous couple of cytotaxomonists, Askell and Doris Love, created a monotypic genus _Yurtsevia_ with one species, _Yurtsevia richardsonii_ (Hook.) A. & D. Love, the segregate of a large collective genus _Anemone_. Boris Alexandrovich Yurtsev died in St. Petersburg on December 14, 2004, after a long fight with cancer. He is survived by his wife Tamara Polozova, daughter Marina, and granddaughter Masha. (This biography is an abbreviated version of the materials sent to BEN by Andrey K. Sytin [astragalus.mail.ru] and Sergey A. Balandin [Balandin@herba.msu.ru].) IN MEMORIAM - BORIS ALEXANDROVICH YURTSEV From: David F. Murray [ffdfm@uaf.edu] I first met Boris at the International Botanical Congress in 1969 (Seattle). Askell Loeve brought us together, saying that Boris was the next Tolmachev, someone I should meet. We chatted briefly and agreed to exchanges of reprints, books, and specimens and eventually of ourselves to the other's country. Reprints and specimens were exchanged immediately, but the exchange of people finally happened in 1981 when Barbara and I visited the Komarov Botanical Institute for the first time. It was a memorable visit, and it set the stage, for me, for the next two decades of study. 1981 was also the first trip for Boris to Alaska, and he published a series of four papers in _Botanicheskij Zhurnal_ of his observations. More trips in the late 1980's led finally to our launching of the _Panarctic Flora Project_. Boris was the primary force behind that initiative, drawing upon his Komarov Botanical Institute workforce and experience gained by the just completed _Arctic Flora of the USSR_ for which he and his co-workers received a State Prize. Boris not only took over editorship of the _Arctic Flora_ after the death of A.I. Tolmachev but also contributed significant portions of Rosaceae and Fabaceae. We joined forces and got the _Panarctic Flora_ project started in the early 1990s, but it was not until we had Scandinavian representation and significant support from Reidar Elven that we really were successful. The annotated checklist is nearly ready for web presentation. Whereas our aim was to provide a unified taxonomy, that has not always been possible, and we have preserved differences, the principal ones being between Boris, the Russian tradition, and Reidar and me. When it came to scientific issues he was a good (stubborn) opponent. In the early 1990s Boris made more trips to Alaska, to the Seward Peninsula under he aegis of the National Park Service Beringian program and one with me to Ellesmere Island as guests of Sylvia Edlund, then of the Geological Survey of Canada. We lived at Tanquary Fiord but Sylvia's work meant we got to helicopter about northernmost Ellesemere, to collect from several sites on the shores of the Arctic Ocean and to revel in the spectacular scenery. Boris was constantly filling notebooks with his observations and impressions. In later life he became a poet and turned out handwritten verse at the drop of a hat, which he often left behind. I recall one on the bulletin board at the Polar Continental Shelf Project operations center at Resolute and we had several in the Herbarium in Fairbanks commemorating collecting trips around Alaska. Ultimately he published a volume of his poems, which, alas, I have not yet seen. He was also a lover of classical music, no surprise for a Russian, and I cherish the times we went to hear choirs or the Philharmonic in St. Petersburg. In the late 1990s Boris was able to become more visible internationally and participated regularly representing Russia at meetings on rare plants (Conservation of Arctic Flora and Fauna), geobotany (Circumpolar Arctic Vegetation Map), and arctic flora. He fought his final illness with the same steadfastness and determination that he botanized. In the field he required very little, mostly he wanted to look at the plants and bemoaned the "permanent shortage of time." He will be missed by his family, his friends, and the world botanical community. PUBLICATION NOTE: ENGLISH TRANSLATION OF THE RUSSIAN ARCTIC FLORA _Flora of the Russian Arctic. A critical review of the vascular plants occurring in the arctic region of the former Soviet Union._ Tolmachev, A. L.; Packer, J. G.;Griffiths, Graham C. D. 3-443-50024-2 English transl. of _Arkticheskaya Flora SSSR: V+VI_ Ed.: Tolmachev, A. L.; Packer, J. G.; Griffiths, Graham C. D. 2000. xxxv+472 p., 5 tables, 166 maps, ISBN 3- 443-50024-2 bound, EUR 127.00 Available from: E.Schweizerbart'sche Verlagsbuchhandlung, Science Publishers, Johannesstr. 3A, D-70176 Stuttgart, Germany E-mail: mail@schweizerbart.de http://www.schweizerbart.de English-language flora of the Russian Arctic (Norwegian Frontier to the Bering Strait). Volume III of the Flora of the Russian Arctic includes treatments of the following nine families: Salicaceae, Betulaceae, Urticaceae, Polygonaceae, Chenopodiaceae, Portulacaceae, Caryophyllaceae, Paeoniaceae and Ranunculaceae. The discussions are illustrated with 166 distribution maps. Once completed, the six volumes of the series will treat about 360 genera, 1650 species and 220 infraspecific taxa. Keys to all genera and species, as well as extensive morphological, ecological and geographical discussions of species characteristic of the Arctic are included. The comprehensive content and accomplished scholarship of the _Flora of the Russian Arctic_ make it an essential part of the libraries of all botanists and botanical institutions concerned with research into northern plants. The Russian Arctic spans 160 degrees of longitude, from the Norwegian frontier to the Bering Strait, thus accounting for about half the circumpolar land mass at arctic latitudes. It is no longer necessary for western botanists to give inadequate consideration to the flora of this vast expanse on account of linguistic difficulties. Flora of the Russian Arctic is a translation of Arkticheskaya Flora SSSR (Flora Arctica URSS), the authoritative work of botanists of the Komarov Botanical Institute prepared under the editorship of A. I. Tolmachev and B. A. Yurtsev. This unabridged translation was prepared by distinguished systematist G. C. D. Griffiths under the editorship of J. G. Packer, Professor Emeritus of Botany at the University of Alberta. Volumes I and II are still available from the University of Alberta Press. See: http://www.uap.ualberta.ca/UAP.asp?LID=41&bookID=90 and http://www.schweizerbart.de/pubs/books/floraofthe-095200001-desc.html ________________________________________________________________ Subscriptions: http://victoria.tc.ca/mailman/listinfo/ben-l Send submissions to aceska@telus.net BEN is archived at http://www.ou.edu/cas/botany-micro/ben/ ________________________________________________________________ From aceska@telus.net Wed Nov 23 16:42:28 2005 From: aceska@telus.net (Adolf Ceska) Date: Wed, 23 Nov 2005 08:42:28 -0800 Subject: [BEN-L]BEN # 353 Message-ID: <000001c5f04c$e85236d0$f1dcb440@HPLAPTOP001> BBBBB EEEEEE NN N ISSN 1188-603X BB B EE NNN N BBBBB EEEEE NN N N BOTANICAL BB B EE NN NN ELECTRONIC BBBBB EEEEEE NN N NEWS No. 352 November 23, 2005 aceska@telus.net Victoria, B.C. ----------------------------------------------------------- Dr. A. Ceska, P.O.Box 8546, Victoria, B.C. Canada V8W 3S2 ----------------------------------------------------------- BEN issues 352 and 353 are dedicated to the memory of BORIS ALEXANDROVICH YURTSEV who died in St. Petersburg on December 14, 2004 BOOK REVIEW: _THE LAST GIANT OF BERINGIA_ From: David F. Murray [ffdfm@uaf.edu] O'Neill, Dan. 2004. _The last giant of Beringia: The mystery of the Bering Land Bridge._ Westview Press, New York, NY. vii+231 p. ISBN 0- 8133-4197-3 [hard cover] Price: US$26.00/CDN$40.00 Available from: Westview Press, 387 Park Avenue South, New York, NY 10016 Tel.: (212) 340-8151, Fax: (212) 340-8115 http://www.westviewpress.com Those who have delved into Beringian history know the name Dave Hopkins. After an illustrious career with the U.S. Geological Survey, mostly working for the Alaska branch, Dave moved to the University of Alaska Fairbanks as a Distinguished Professor of Quaternary Science. At UAF he was the magnet that attracted faculty and students-botanists, zoologists, geologists, anthropologists, oceanographers-- and the glue that held them together through the activities of the Alaska Quaternary Center. Dan O'Neill has written a biography of Dave, _The last giant of Beringia_, the giant referring to Dave, but clearly with a play on the word synonymous with Pleistocene megafauna. Dan has provided background on Dave's upbringing and early years in New Hampshire (born in 1921), the early development of his broad interests in natural history, his education, and on his long and productive professional career of more than 50 years, most of it in Alaska. In recent correspondence I have had with others who worked more closely with Dave and knew him better, I learned that we had each found something in the Dan's account we hadn't known before. Dan hit the high points of the Bering Land Bridge, its history and impact on our biota, when during glacial intervals Alaska was biologically more an extension of northeast Asia than a part of North America. He described how Dave began to investigate Beringian history, step by step, working with fossil mollusks, sea floor morphology, ancient shorelines, paleosols, beetles, pollen and macrofossils, and the artifacts of early humans. He showed how the results of one study led to another, additively, until a coherent thread had been established. Dave was an intellectual omnivore, the quintessential interdisciplinary scientist, who could seize new ideas from any and all quarters, but when any one of them failed the test, happily discarded it and eagerly move on. Several important players have smaller parts in the book, and Dan has done justice to the critical influences of botanists, Eric Hult,n and Bob Sigafoos, one archaeologist in particular, Louis Giddings, and many, many others who contributed hugely to Dave's understanding of Beringian paleoecology. Dave was esteemed by his Russian colleagues just as has Dan described. I took reprints and greetings from Dave to Moscow on one of my trips and met with his old friends who made it abundantly clear that Dave was truly a good friend. Boris Yurtsev was one of those Russian friends. On a visit to Fairbanks, Boris was able to join the annual Quaternary field trip that Dave had instigated as a regular early fall event. We tented overnight on the "green" at Central and then went out to look at Quaternary gravels at local placer mines. Fall color was at its peak and the aspen and birch covered hillsides were a blaze of gold and Boris enchanted. Dave's work led to symposium volumes to which he contributed papers or ones for which he was the motivating force, the most important of these being the _The Bering Land Bridge_ in 1967 and _Paleoecology of Beringia_ in 1982. More recently in 2001 there appeared a festschrift to Dave, _Beringian Paleoenvironments_, which is another important synthesis. It was the outcome of a conference that included a very moving evening with Dave, some slides, and his very personal reminiscences. Boris Yurtsev was there with a large Russian contingent, and the paper Boris published in the festschrift is the most complete statement we have of his tundra-steppe concept. Dave Hopkins died in 2001. Dave's legacy is his family and friends, seminal papers, and lots of colleagues and former students, themselves now important figures in Quaternary science. He was a giant. PROCEEDINGS FROM THE PANARCTIC FLORA 1998 SYMPOSIUM Nordal, I. & V. Yu. Razzhivin [eds.] 1999. _The species concept in the high north a Panarctic flora initiative._ Det norske videnskaps-akademi. Skrifter. 1. Mat.- naturv. klasse. Ny serie (nr. 38). Oslo. 418 p. ISBN 82-90888-33-3; ISSN 1502- 0096 [soft cover] Price: US$40.00 Limited number of copies is available from: Inger Nordal [inger.nordal@bio.uio.no] Thirty-two contributions by US, Canadian, Icelandic, Norwegian, Swedish and Russian botanist deal with problems related to compiling the Panarctic Flora. This Flora would include all the Arctic regions. The "Panarctic Year" at the Norwegian Centre of Advanced studies was initiated by an international symposium, September 28 to October 3, 1998, with about 30 participants. Section I in this volume includes the proceedings from this symposium. With the extant knowledge of the history, palaeogeography, evolution, and taxonomy of the Arctic, the editors wanted to be open to different views, and contrasting, even conflicting, views might be found. During the "Panarctic Year" several workshops (or the so-called "expert group meetings") have been organised on critical genera/families with complicated taxonomy, obviously treated differently within Russian, Nordic, and/or American tradition. The reports from these workshops are presented in Section II and include notes on _Cerastium_, _Draba_, _Dryas_, _Papaver_, Poaceae, _Potentilla_, _Salix_, and _Saxifraga_. Section III contains reports on an earlier meeting of the Panarctic Flora committee, zonal subdivision of the Arctic, and an example of a checklist of Arctic members of the Saliaceae family. A COMPILATION OF ADDITIONS TO THE FLORA OF THE CONTINENTAL PORTIONS OF NORTHWEST TERRITORIES AND NUNAVUT From: P. M. Catling, W. J. Cody and G. Mitrow, Biodiversity, National Program on Environmental Health, Agriculture and Agri-food Canada, Wm. Saunders Bldg., Central Experimental Farm, Ottawa, Ontario, Canada K1A 0C6 E-mail: [catlingp@agr.gc.ca] Introduction The protection of biodiversity in the Canadian north is receiving increasing attention as settlement and exploitation of natural resources expand. Invasive species are rapidly entering the region (Wein et al. 1992, Cody et al. 2000) which includes many native species of concern (Cody 1979, McJannet et al. 1995, Talbot et al. 1999) and many that are important to native people and wildlife (e.g. Marles et al. 2000). A strong and ever growing commitment to the protection of biodiversity is evident in territorial initiatives such as the Northwest Territories Biodiversity Action Plan (NWT Biodiversity Team 2004). With the increasing interest in the protection of natural resources there is an increasing need for accurate information on the flora including particularly the information needed for plant identification. There is also a great international interest in the Canadian portion of the circumpolar flora as a result of cooperative work on classification and evolution of circumpolar plants involving all circumpolar countries. This work also requires that information be current and accessible. The information presented here responds to these needs. The list includes species that are new to the continental areas of Northwest Territories and Nunavut together, and thus new to the classic flora (Porsild and Cody 1980) that covers this region. The area In 1980 when the manual of _Vascular Plants of the Continental Northwest Territories, Canada_ (Porsild & Cody 1980) was published the "Territories" included a continental (excluding oceanic islands) land and freshwater area of approximately 3/4 of a million square miles. It extended from the Yukon border north of 60 degrees east to Hudson Bay and was bounded on the north by the Arctic Ocean. On April 1, 1999, the eastern portion of this territory (including all of the District of Keewatin and much of the District of Mackenzie and the District of Franklin) became a part of a separate territory called Nunavut which has a continental area of 350,000 square miles. Recent work Since 1980 there have been several regional studies of the flora and vegetation of mainland parts of both Northwest Territories (NT) and Nunavut (NU). Some of these have emphasized phytogeography and included reference to the discovery of additional species ( e.g. Cody et al. 2003) while others have not, often being concerned primarily with detailed information on vegetation composition (e.g. Cody et al. 1984). In addition numerous systematic studies of particular groups of plants have been published and many of these contain additional plant records, i.e. species additional to those included in the manual. To use the guide effectively an update to the information is required. Here we provide the update with an indication of sources of additional information. Since this compilation was prepared two additional sources have become available but the results have not been included here (see Catling 2005 and Catling 2006). Methods This compilation includes all additions to the Porsild and Cody (1980) manual known to us and is also based on (1) a literature search in Biosis using such key words as "addition", "Northwest Territories", "new record", "new plant", "flora;" (2) an analysis of the maps in "Flora of the Canadian Arctic Archipelago" (Aiken et al. 2002), (3) a search for additional records in the recently published volumes of Flora of North America (including volumes 2, 3, 4, 22, 23, 25, and 26; (3) a survey of monographs published in Rhodora; and (4) a survey of articles in The Canadian Field-Naturalist. We suspect that with this work we have compiled over 95% of the additions. We will appreciate learning of any that we have overlooked. This compilation applies only to the combined continental area of NT and NU, i.e. continental N.W.T. in the sense of Porsild and Cody (1980). The northernmost peninsulas of Nunavut, the Boothia Peninsula and the Melville Peininsula, are traditionally not included in the continental area (Porsild & Cody, 1980, p. 2). Additional taxa may include species, subspecies, varieties, forms, hybrids and differentiated races potentially worthy of taxonomic recognition. Additions may result from new discoveries in the area, corrections to existing reports or newly defined taxa. The taxa are arranged by family and this will help to ensure that users of the 1980 flora are aware of more recently discovered taxa related to those that can be identified using the the 1980 text. More information on these species is available in the references provided. Synopsis Porsild and Cody (1980) listed 1113 species. The published additions of 111 taxa listed here includes 94 species, 9 infraspecific taxa (i.e. additional subspecies, varieties and forms of species listed by Porsild & Cody in 1980) and 8 hybrids. Based on the recorded additions and the deletions of 6 species (listed below), 1201 species are known from the combined continental areas of NT and NU. An accurate number of taxa (all categories including different subspecies, varieties, forms and hybrids) is not available at present. Of the additional species, 68 are native and 26 are introduced. The families with the largest representation among the additional species are the grasses (Poaceae) and sedges (Cyperaceae). Additions to Northwest Territories (NT) and Nunavut (NU) * = introduced ISOETACEAE _Isoetes lacustris_ L. (_I. macrospora_Dur.; Cody 1996). EQUISETACEAE _Equisetum_ X _litorale_ Kuehlewein ex Ruprecht (_E. arvense_ L. X _E. fluviatile_ L.; Hauke 1993). OPHIOGLOSSACEAE _Botrychium simplex_ E. Hitchcock (McJannet et al. 1995, Cody 1996). _Botrychium spathulatum_ W. H. Wagner (Wagner & Wagner 1993, Cody 1996). POLYPODIACEAE _Polypodium sibiricum_ Siplivinsky. Most of the _Polypodium_ plants in NT & NU are referable to this species rather than the previously reported _Polypodium virginianum_ L. which is known only from an isolated area near the southern border (Haufler et al. 1993). PINACEAE _Abies bifolia_ A. Murray (Thieret 1993). ACORACEAE _Acorus americanus_ (Raf.) Raf. Previously reported as _A. calamus_ L. which is now treated as a separate species with a more southern distribution (Thompson 2000). LEMNACEAE _Lemna turionifera_ Landolt. _Lemna minor_ L. has been split into _L. minor_ and _L. turionifera_. Plants reported as the former from NT are now recognized as _Lemna turionifera_ (Landolt 2000). SPARGANIACEAE _Sparganium emersum_ Rehmann (Kaul 2000). POTAMOGETONACEAE _Potamogeton obtusifolius_ Mertens & W. D. J. Koch (Cody 1996). _Potamogeton pusillus_ L. subsp. _tenuissimus_ (Mertens & W.D.J. Koch) R.R. Haynes & Hellquist (Haynes & Hellquist 2000). _Stuckenia filiformis_ (Pers.) Boerner subsp. _filiformis_. Plants reported previously under this name were placed with the _P. filiformis_ Pers. var. _borealis_ (Raf.) St. John, now _Stuckenia filiformis_ subsp. _alpina_ (Blytt) R.R. Haynes, Les, & Kral. Thus _Stuckenia filiformis_ subsp. _filiformis_ is an addition (Haynes & Hellquist 2000). _Stuckenia filiformis_ (Persoon) Boerner subsp. _occidentalis_ (J.W. Robbins) R.R. Haynes, Les, & M. Kral (Haynes & Hellquist 2000). _Zannichellia palustris_ L. (Cody 1998). ALISMATACEAE _Alisma plantago-aquatica_ L. var. _americana_ J. A. Schultes (_A. triviale_ Pursh; Cody 1996). POACEAE *_Agrostis stolonifera_ L. (Cody et al. 2000). *_Alopecurus arundinaceus_ Sobol (Cody et al. 2000). *_Alopecurus pratensis_ L. (Cody et al. 2000). *_Bromus commutatus_ Schrader (Cody et al. 2000). *_Bromus hordeaceus_ L. (_B. mollis_ auct. non L.; Cody et al. 2000). *_Bromus squarrosus_ L. (Cody et al. 2000). _Calamagrostis holmii_ Lange (McJannet et al. 1995). _Danthonia spicata_ (L.) Beauvois ex Roemer & J. A. Schultes (Cody 1996, Cody, Reading & Line 2003). *_Deschampsia elongata_ (Hook.) Munro (Cody, Reading & Line 2003). _Festuca hyperborea_ Holmen ex Frederiksen. Shown to occur in Nunavut by Aiken & Darbyshire (1990) and in Northwest Territories by Aiken et al. (2002). *_Festuca trachyphylla_ (Hackel) Krajina (Cody et al. 2000). *_Festuca valesiaca_ Schleicher ex Gaudin subsp. _sulcata_ (Hackel) Schinz & R. Keller (Cody et al. 2000). *_Lolium perenne_ L. subsp. _multiflorum_ (Lamarck) Husnot (Cody et al. 2000). *_Lolium perenne_ L. subsp. _perenne_ (Cody et al. 2000). _Pleuropogon sabinei_ R. Brown (Cody et al. 1989, also Korol 1992). *_Poa annua_ L. (Cody et al. 2000). _Poa hartzii_ Gandoger Arctic Ocean coastline (Gillespie et al. 1997, Gillespie & Boles 2001, Aiken et al. 2002). _Poa pseudoabbreviata_ Roshevitz (Cody 1996). *_Secale cereale_ L. (Cody et al. 2000). *_Sorghum halepense_ (L.) Pers. (Barkworth 2003). *_Triticum aestivum_ L. (Cody et al. 2000). *_Vulpia bromoides_ (L.) S. F. Gray (Cody et al. 2000). CYPERACEAE _Carex bigelowii_ Torrey ex Schweinitz X _subspathacea_ Wormskj. Possibly refers to the continental area (Cayouette & Catling 1992). _Carex bigelowii_ Torrey ex Schweinitz subsp. _lugens_ (T. Holm) T. V. Egorova. Included in _C. bigelowii_ by Porsild and Cody (1980, see Standley et al. 2002). _Carex flava_ L. (Crins 2002). _Carex hoodii_ Boott (Cody, Reading & Line 2003). _Carex microptera_ Mack. (_Carex festivella_ Mack. which was included in _C. macloviana_ d'Urv. by Porsild & Cody (1980), see Cody, Reading & Line (2003) and Mastrogiuseppe et al. (2002). _Carex petasata_ Dewey (Cody, Reading & Line 2003). _Carex prairea_ Dewey (Cody 1996). _Carex sterilis_ Willdenow (Reznicek 2002). _Carex utriculata_ Boott in W.J. Hook.. Previously often included with _C. rostrata_ Stokes (Reznicek & Ford 2002). _Eriophorum tenellum_ Nuttall (Ball & Wujek 2002). _Eriophorum_ X _medium_ Andersson subsp. _album_ J. Cayouette (_E. russeolum_ Fries ex Hartman subsp. _leiocarpum_ X _E. scheucheri_ Hoppe subsp. _scheuchzeri_; Cayouette 2004). _Schoenoplectus acutus_ (Muhlenberg ex Bigelow) A. L"ve & D. L"ve var. _acutus_ (Smith 2002). JUNCACEAE _Juncus mertensianus_ Bongard (Brooks & Clemants 2000). _Juncus tenuis_ Willdenow (Brooks & Clemants 2000). LILIACEAE *_Allium fistulosum_ L. (Cody 1996). _Maianthemum racemosum_ (L.) Link subsp. _amplexicaule_ (Nutt.) LaFrankie (_Smilacina amplexicaulis_ Nutt.; Lafrankie 2002). IRIDACEAE _Sisyrinchium septentrionale_ E.P. Bicknell (Cholewa & Henderson 2002). ORCHIDACEAE _Cypripedium parviflorum_ var. _makasin_ (Farwell) Sheviak (_C. pubescens_ var. _makasin_ Farwell; Sheviak 2002). _Malaxis monophyllos_ L. Swartz var. _brachypoda_ (A. Gray) Morris & Eames (_M. brachypoda_ [Gray] Fern.; Cody & Johnston 2003). SALICACEAE _Salix arctica_ Pallas X _S. glauca_ L. (Cody & Reading 2005). _Salix raupii_ Argus (Argus 1986). _Salix rotundifolia_ Trautvetter subsp. _rotundifolia_ (Cody, Reading & Line 2003, also Cody & Reading 2005). _Salix sphenophylla_ Skvortzov (McJannet et al. 1995). BETULACEAE _Alnus viridis_ (Villars) DeCandolle subsp. _fruticosa_ (Ruprecht) Nyman (Furlow 1997). _Alnus viridis_ (Villars) Lamarck & DeCandolle subsp. _sinuata_ (Regel) Loeve & Loeve (Furlow 1979, 1997). _Betula nana_ L. subsp. _exilis_ (Sukaczev) Hult. Previously combined with _B. glandulosa_ Michx. in Porsild & Cody 1980 (Furlow 1997). _Betula_ X _uliginosa_ Dugle (_B. glandulosa_ Michx. X _B. neoalaskana_ Sarg.; Jasieniuk & Johnson 1979). _Betula_ X _eastwoodiae_ Sarg. (_B. glandulosa_ Michx. X _B. occidentalis_ Hook.; Furlow 1997). _Betula_ X _sargentii_ Dugle (_B. glandulosa_ Michx. X _B. pumila_ L.; Jasieniuk & Johnson 1979). POLYGONACEAE *_Polygonum fowleri_ Robinson (Cody, Reading & Line 2003). _Polygonum ramosissimum_ Michaux (Cody, Reading & Line 2003). CHENOPODIACEAE _Chenopodium simplex_ (Torrey) Rafinesque (_C. hybridum_ L. var. _simplex_ Torr., _C. gigantospermum_ Aellen; Clemants & Mosyakin 2003). *_Corispemum villosum_ Rydb. (Cody, Reading & Line 2003). _Corispermum hookeri_ Mosyakin (Cody, Reading & Line 2003). _Corispermum ochotense_ Ignatov (Cody, Reading & Line 2003). *_Corispermum orientale_ Lamarck var. _emarginatum_ (Rydb.) Macbride (Cody et al. 2000). CARYOPHYLLACEAE _Sagina procumbens_ L. (Rabeler & Thieret 1997). _Silene sorensenis_ (Boivin) Bocquet. Shown to occur in the Mackenzie delta region by Aiken et al. (2002). _Silene uralensis_ (Ruprecht) Bocquet subsp. _ogilviensis_ (A.E. Porsild) Brunton (Cody, Reading & Line 2003). RANUNCULACEAE _Anemone multifida_ Poiret var. _saxicola_ Boivin (Dutton et al. 1997). _Anemone patens_ L. var. _multifida_ Pritzel (Dutton et al. 1997). _Ranunculus aquatilis_ L. var. _aquatilis_ (Whittemore 1997). _Ranunculus flammula_ L. var. _ovalis_ (J.M. Bigelow) L.D. Benson (Whittemore 1997). _Ranunculus_ X _spitzbergensis_ (Nathorst) Hadac (_R. lapponicus_ L. x _R. pallasii_ Schlecht.; Cody et al. 1988). _Ranunculus gmelinii_ DC. X _R. hyperboreus_ Rottb. ( _R. yukonensis_ Britt., _R. hyperboreus_ Rottb. var. _turquetilianus_ Polunin; Cayouette et al. 1997). PAPAVERACEAE _Papaver radicatum_ Rottb. subsp. _polare_ Tolmachew (Kiger & Murray 1997). BRASSICACEAE _Arabis calderi_ G. Mulligan (Mulligan 1995). _Arabis holboellii_ Hornemann var. _secunda_ (Howell) Jepson (Cody 1996). Also sometimes included in the genus _Boechera_ (Loeve & Loeve 1976, Aiken et al. 2002) as _Boechera holboellii_ (Hornem.) Loeve & Loeve but some recent authors have retained the genus name _Arabis_ (Mulligan 1995, Rollins 1997). _Arabidopsis lyrata_ (L.) O'Kane and Al-Shehbaz subsp. _lyrata_. Reported as _Arabis lyrata_ L. by Cody (1996) who used the name in the strict sense, i.e. not including _A. lyrata_ var. _kamchatica_ Fischer and _A. kamchatica_ (Fischer) Ledeb. Mulligan (1995) treated the latter as a species while O'Kane and Al-Shehbaz (1997) accepted it as a subspecies. _Arabis pinetorum_ Tidestrom (Cody 1996). _Braya pilosa_ Hook. This plant of sandy sea shore is apparently not a synonym of B. glabella of damp interior tundra as postulated Porsild & Cody 1980 (McJannet et al. 1995, Aiken et al. 2002). It was treated as a species by Rollins (1993) who noted that it is "known only from the type locality at the mouth of the Mackenzie River. *_Descurainia incisa_ (Engelmann ex Gray) Britton subsp. _incisa_ (_D. richardsonii_ O.E. Schultz subsp. _incisa_ (Engelm.) Detling; Cody 1996, Cody, Reading & Line 2003). _Draba oblongata_ R.Br. ex DC. was expected to be discovered on the mainland coast by Porsild & Cody (1980) and recently mapped from the base of the Boothia Peninsula by Aiken et al. (2002). Several species (including _Draba subcapitata_ Simmons and _Epilobium arcticum_ Samuelsson) are widespread on the arctic islands but known on the mainland only from the Boothia Peninsula or from there and a very few other coastal sites. _Draba stenoloba_ Ledeb. (Cody, Reading & Line 2003). _Draba subcapitata_ Simmons (Cody et al. 1992). _Erysimum coarctatum_ Fernald (Cody, Reading & Line 2003). CRASSULACEAE *_Sedum spurium_ Bieberstein (Wein et al. 1992). ROSACEAE _Potentilla hyparctica_ Malte. A northern race, possibly worthy of taxonomic recognition, has been differentiated by Aiken et al. (2002). It occurs in the mainland area at the base of the Boothia Peninsula and on the coast south of King William Island. FABACEAE _Astragalus eucosmus_ Robinson f. _albinus_ Fernald (Cody & Reading 2005). *_Caragana arborescens_ Lamarck (Wein et al. 1992). *_Medicago sativa_ L. subsp. _falcata_ (L.) Arcang. (Wein et al. 1992, Cody, Reading & Line 2003). *_Vicia cracca_ L. (Wein et al. 1992). VIOLACEAE _Viola selkirkii_ Pursh ex Goldie (Cody 1996). ONAGRACEAE _Epilobium hornemanii_ Reichenb. (Cody 1996). _Epilobium latifolium_ L. f. _albiflorum_ Nathorst (Cody & Reading 2005).This species is often placed in the genus _Chamerion_ as _Chamerion latifolium_ (L.) Holub. APIACEAE _Podistera macounii_ (Coulter & Rose) Mathias & Constance (_Ligusticum mutellinoides_ (Crantz) Willar; McJannet et al. 1995). ERICACEAE _Vaccinium ovalifolium_ Sm. (Cody, Reading & Line 2003). GENTIANACEAE _Halenia deflexa_ (J. E. Smith) Grisebach (Cody, Reading & Line 2003). POLEMONIACEAE _Polemonium boreale_ Adams f. _albiflorum_ Cody (Cody, Reading & Line 2003). LAMIACEAE _Monarda fistulosa_ L. var. _menthifolia _(Graham) Fernald (Straley 1986). _Physostegia ledinghamii_ (Boivin) Cantino. Loan's collection from Salt River at DAO is a paratype. Reported by Porsild & Cody (1980) as _P. parviflora_ Nutt. ex Gray (Cantino 1981, Feswick & Catling 2004). SCROPHULARIACEAE *_Veronica longifolia_ L. (Cody 1996). ASTERACEAE *_Crepis tectorum_ L. (Cody, Reading & Line 2003). _Packera ogotorukensis_ (Packer) Loeve & Loeve. Reported as _Senecio ogotorukensis_ Packer (Cody 1996). Notes _Carex foenea_ Willd. A misapplied name in Porsild and Cody (1980), now _Carex siccata_ Dewey based on examination of numerous specimens at DAO herbarium supporting its widespread occurrence in southern NT. _Carex paleacea_ Schreb. ex Wahlenb. The map in Standley et al. (2002) is incorrect in indicating the presence of this species in the area (J. Cayouette pers. comm.). _Corispermum hyssopifolium_ L. Deleted by Cody, Reading & Line (2003). _Danthonia intermedia_ Vasey. Revised to _D. spicata_ (L.) Beauv. ex Roem. & Schult. (Cody 1996, Cody, Reading & Line 2003). _Eleocharis compressa_ Sullivant. The report of _E. compressa_ in Porsild & Cody (1980) is apparently based on 2 sheets at DAO from the vicinity of Heart Lake Biological Station 60 deg. 51'N, 116 deg. 37'W, collected in 1972 by S. S. Talbolt (3742 and 2310). Since the translucent apex of scales is less than 0.5 mm long, and the culms are 2 times wider than thick, these specimens seem best placed with _E. elliptica_ Kunth. _Eleocharis compressa_ was not reported from NT by Smith et al. (2002) and _E. elliptica_ is a new record listed above. _Erysimum inconspicuum_ (S. Wats.) MacM. Revised to _E. coarctatum_ Fern. (Cody, Reading & Line 2003). _Nymphaea tetragona_ Georgi subsp. _leibergii_ (Morong) Porsild. Now treated as _Nymphaea leibergii_ Morong but the material reported under the former name by Porsild & Cody (1980) is referable to _Nymphaea tetragona_ Georgi. _Physostegia parviflora_ Nutt. ex Gray. This species reported by Porsild and Cody (1980) is to be deleted, the specimens being referable to _P. ledinghamii_ (Boivin) Cantino (Cantino 1981, Feswick & Catling 2004). _Saxifraga tenuis_ (Wahlenb.) H.Sm. ex Lindm. Deleted by Cody (1996) on the basis if revision of specimens reported by Korol (1992) to _S. nivalis_ L. Acknowledgement Helpful comments were provided by Dr. Jacques Cayouette. Literature Cited Aiken, S.G., M.J. Dallwitz, L.L. Consaul, C.L. McJannet, L.J. Gillespie, R.L. Boles, G.W. Argus, J.M. Gillett, P.J. Scott, R. Elven, M.C. LeBlanc, A.E. Zamluk, and A.K. Brysting. 2002. _Flora of the Canadian Arctic Archipelago: Descriptions, Illustrations, Identification, and Information Retrieval._ Version: 25th March 2002. http://www.mun.ca/biology/delta/arcticf/ Aiken, S.G. and S. Darbyshire. 1990. _Fescue grasses of Canada._ Agriculture Canada Publication 1844/E. 113 p. Argus, G. W. 1986. _Salix raupii_, Raup's Willow, new to the Flora of Alberta and the Northwest Territories. _Canadian Field-Naturalist_ 100 (3): 386-388. Ball P. W. & D. E. Wujek 2002. 2. _Eriophorum_ L.. Pp. 21-27 in Flora of North America Editorial Committee, eds. _Flora of North America Volume 23, Magnoliophyta: Commelinidae, (in part): Cyperaceae._ Oxford University Press, New York. Barkworth, M. E.2003. 26.08 _Sorghum_ Moench. Pp. 626-630 in Barkworth M.E., K.M. Kapels, S. Long & M. B. Piep, eds. _Flora of North America Volume 25, Magnoliophyta: Commelinidae (in part: Poaceae, part 2)._ Oxford University Press, New York. Brooks R. E. & S. E. Clemants 2000. 209. Juncaceae Jussieu Pp. 211-268 in Flora of North America Editorial Editorial Committee, eds. _Flora of North America Volume 22, Magnoliophyta: Alismatidae, Arecidae, Commelinidae (in part), and Zingiberidae._ Oxford University Press, New York. Cantino, P. D. 1981. Change of status for _Physostegia virginiana_ var. ledinghamii (Labiatae) and evidence for a hybrid origin. _Rhodora_ 83 (833): 111-118. Catling, P.M. 2005. Identification and distribution of the small white water-lilies, _Nymphaea tetragona_ and _N. leibergii_, in Northwest Territories. _BEN - Botanical Electronic News_, 348. Catling, P. 2006. Additions to the flora of the continental Northwest Territories from the Great Slave area. _Canadian Field-Naturalist_. [in press] Cayouette, J. 2004. A taxonomic review of the _Eriophorum russeolum_ - _E. scheuchzeri_ complex (Cyperaceae) in North America. _Sida_ 21(2): 791-814. Cayouette, J., M. Blondeau, and P. M. Catling. 1997. 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Elvebakk. 1999. _Atlas of the rare endemic vascular plants of the arctic._ Conservation of Arctic Flora and Fauna (CAFF) Technical Report Number 3. U.S. Fish and Wildlife Service, Anchorage, Alaska, 99503, iv + 73 p. Thieret J. W. 1993. 3. Pinaceae Lindley Pp. 352-398 in Flora of North America Editorial Editorial Committee, eds. _Flora of North America Volume 2, Pteridophytes and Gymnosperms._ Oxford University Press, New York. Thompson, S. A. 2000. 202. ACORACEAE C. Agardh. Pp. 124-127 in Flora of North America Editorial Editorial Committee, eds. _Flora of North America Volume 22, Magnoliophyta: Alismatidae, Arecidae, Commelinidae (in part), and Zingiberidae._ Oxford University Press, New York. Wagner W. H., & F. S. Wagner 1993. 6. Ophioglossaceae C. Agardh Pp. 85- 106 in Flora of North America Editorial Editorial Committee, eds. _Flora of North America Volume 2, Pteridophytes and Gymnosperms._ Oxford University Press, New York. Wein, R. W., G. Wein, S. Bahret, and W. J. Cody. 1992. Northward invading non-native vascular plant species in and adjacent to Wood Buffalo National Park, Canada. _Canadian Field- Naturalist_ 106 (2): 216- 224. Whittemore A.T. 1997. 2. _Ranunculus_ L. Pp. 88-135 in Flora of North America Editorial Committee, eds. _Flora of North America Volume 3, Magnoliophyta: Magnoliidae and Hammamelidae._ Oxford University Press, New York. ________________________________________________________________ Subscriptions: http://victoria.tc.ca/mailman/listinfo/ben-l Send submissions to aceska@telus.net BEN is archived at http://www.ou.edu/cas/botany-micro/ben/ ________________________________________________________________